Athleta

Athleta Temporal range: Cretaceous to Recent
Athleta ficulina, a fossil species from Miocene
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
Subclass:	Caenogastropoda
Order:	Neogastropoda
Superfamily:	Volutoidea
Family:	Volutidae
Genus:	Athleta Conrad, 1853[1]
Type species
† Voluta rarispina Lamarck, 1811
Synonyms[2]
Athleta (Athleta) Conrad, 1853· accepted, alternate representation Athleta (Neoathleta) Sacco, 1890 †· accepted, alternate representation Athleta (Oligoathleta) Lozouet, 2019 †· accepted, alternate representation Athleta (Ternivoluta) Martens, 1897· accepted, alternate representation Athleta (Volutocorbis) Dall, 1890 †· accepted, alternate representation Athleta (Volutospina) Newton, 1906 †· accepted, alternate representation Notoplejona Marwick, 1926 † Voluta (Ternivoluta) Martens, 1897 Volutilithes (Neoathleta) Sacco, 1890 † Volutilithes (Volutocorbis) Dall, 1890 (original rank) Volutocorbis Dall, 1890 Volutospina Newton, 1906 (original rank) Volutospina (Athleta) Conrad, 1853 †

Athleta is a genus of sea snails, marine gastropod mollusks in the family Volutidae.^[2]

Species within the genus Athleta include:

• † Athleta affinis (Brocchi, 1814)
• Athleta boswellae (Rehder, 1969)^[3]
• † Athleta delvallei Astibia, Merle & Pacaud, 2018
• Athleta disparilis (Rehder, 1969)^[4]
• Athleta easoni Petuch & Berschauer, 2017
• Athleta epigona (Martens, 1904)^[5]
• † Athleta ficulina (Lamarck, 1811)
• † Athleta gabriellis (Benoist, 1874)
• Athleta gilchristi (G.B. Sowerby III, 1902)^[6]
• Athleta glabrata (Kilburn, 1971)^[7]
• Athleta insperata Darragh, 1979^[8]
• Athleta kilburni (Rehder, 1974)^[9]
• † Athleta lata (Marwick, 1926)
• Athleta lutosa (Koch, 1948)^[10]
• † Athleta maculosus Lozouet, 2019
• Athleta magister (Kilburn, 1980)
• † Athleta marwicki P. A. Maxwell, 2003
• † Athleta memoirae Lozouet, 2019
• † Athleta mimica P. A. Maxwell, 2003
• Athleta mozambicana (Rehder, 1972)^[11]
• Athleta nana (Rehder & Weaver, 1974)^[12]
• † Athleta necopinata Suter, 1917
• †Athleta permulticostatus (Telegdi-Roth, 1914)
• † Athleta peyreirensis Lozouet, 2019
• Athleta pisororum H. Morrison, 2006
• † Athleta rathieri (Hébert, 1849)
• Athleta rosavittoriae (Rehder, 1981)^[13]
• Athleta semirugata (Rehder & Weaver, 1974)^[14]
• Athleta pisororum Morrison, 2006^[15]
• † Athleta spinosa (Linnaeus, 1758)
• Athleta studeri (Martens, 1897)^[16]
• † Athleta subaffinis (d'Orbigny, 1852)
• † Athleta subambiguus (d'Orbigny, 1852)
• † Athleta subelegans (d'Orbigny, 1852)
• Athleta taikoensis P. A. Maxwell, 2003
• † Athleta telegdyi (Gaál, 1938)
• † Athleta tournoueri (Peyrot, 1928)

subgenus Athleta Conrad, 1853

subgenus Ternivoluta Martens, 1897^[17]

This article is missing information about Parentheses in this section "Fossil species" may be missing or incorrect.. Please expand the article to include this information. Further details may exist on the talk page. (January 2018)

subgenus Athleta Conrad, 1853^[18]

• † Athleta fayiumensis Abbass, 1967^[18]
• † Athleta strombiformis Deshayes, 1835^[18]

subgenus † Neoathleta Bellardi, 1890^[18]

• † Athleta barrandii Deshayes, 1865^[18]
• † Athleta cithara Lamarck, 1802^[18]
• † Athleta geminata Edwards, 1854^[18]
• † Athleta labrellus Lamarck, 1802^[18]
• † Athleta mutatus Deshayes, 1835^[18]

subgenus Volutocorbis Dall, 1890^[18]

• † Athleta burtoni Vredenburg, 1923^[18]
• † Athleta eugeniae Vredenburg, 1923^[18]
• † Athleta lasharii Merle & Pacaud, 2014^[18]
• † Athleta wynnei Cox, 1930^[18]

Sometimes the following species used to be recognized in the separate genus:

• † Volutocorbis humei Abbass, 1967^[18]
• † Volutocorbis sohli Abbass, 1967^[18]
• † Volutocorbis crenulifera Bayan, 1870^[18]
• † Volutocorbis digitalina Lamarck, 1810^[18]
• † Volutocorbis elevata Sowerby, 1840^[18]
• † Volutocorbis inornatus Oppenheim, 1906^[18]
• † Volutocorbis lima Sowerby, 1823^[18]
• † Volutocorbis minutus Perrilliat et al., 2006^[18]
• † Volutocorbis pakistanica Eames, 1952^[18]
• † Volutocorbis scabricula Linnaeus, 1758^[18]

subgenus † Volutopupa Dall, 1890^[18]

• † Athleta citharopsis Merle & Pacaud, 2014^[18]
• † Athleta intercrenatus Cossmann & Pissarro, 1909^[18]

subgenus Volutospina Newton, 1906^[18]

• † Athleta ambiguus Solander in Brander, 1766^[18]
• † Athleta athleta Solander in Brander, 1877^[18]
• † Athleta dentata Sowerby, 1839^[18]
• † Athleta depauperatus Sowerby, 1823^[18]
• † Athleta jacobsi Vredenburg, 1921^[18]
• † Athleta luctator Sowerby, 1823^[18]
• † Athleta mekranica Vredenburg, 1925^[18]
• † Athleta nodosa Sowerby, 1829^[18]
• † Athleta noetlingi Cossmann & Pissarro, 1909^[18]
• † Athleta sindiensis Vredenburg, 1925^[18]
• † Athleta spinosa Linnaeus, 1758^[18]
• † Athleta subambigua d'Orbigny, 1852^[18]
• † Athleta suspensus Solander in Brander, 1766^[18]
• † Volutospina conicoturrita Newton, 1922^[18]
• † Volutospina kohatica Eames, 1952^[18]
• † Volutospina multispinosa Newton, 1922^[18]

subgenus ?

• † Athleta clayi^[18]
• † Athleta daviesi Cox, 1930^[18]
• † Athleta ficulina Lamarck, 1811^[18]
• † Athleta haleanus^[18]
• † Athleta lisbonensis^[18]
  • † Athleta lisbonensis forma crockettensis Plummer, 1933^[18]
• † Athleta lugardi Newton, 1922^[18]
• † Athleta petrosa Conrad, 1833^[18]
• † Athleta pygmaea Bellardi, 1890^[18]
• † Athleta rarispina Lamarck, 1811^[18]
• † Athleta sayanus Conrad, 1833^[18]
• † Athleta symmetricus Conrad, 1854^[18]
• † Athleta wynnei Cox, 1930^[18]

• Athleta abyssicola (Adams & Reeve, 1848):^[19] synonym of Capensisvoluta abyssicola (A. Adams & Reeve, 1848) (superseded combination)
• Athleta anticingulata (F. McCoy, 1866) †: synonym of Athleta anticingulata anticingulata (F. McCoy, 1866) †: synonym of Ternivoluta anticingulata anticingulata (McCoy, 1866) †
• Athleta antiscalaris (F. McCoy, 1866) †: synonym of Ternivoluta antiscalaris (F. McCoy, 1866) †
• Athleta antispinosa (Tate, 1899) †: synonym of Ternivoluta antispinosa (Tate, 1899) † (superseded combination)
• Athleta boswellae (Rehder, 1969): synonym of Capensisvoluta boswellae (Rehder, 1969) ( superseded combination)
• Athleta bungae Darragh, 1971 †: synonym of Ternivoluta bungae (Darragh, 1971) † (superseded combination)
• Athleta curvicostata Darragh, 1971 †: synonym of Ternivoluta curvicostata (Darragh, 1971) † (superseded combination)
• Athleta disparilis (Rehder, 1969): synonym of Capensisvoluta disparilis (Rehder, 1969) (superseded combination)
• Athleta easoni Petuch & Berschauer, 2017: synonym of Capensisvoluta easoni (Petuch & Berschauer, 2017) (original combination)
• Athleta emmanuelae (Rosso, 1985): synonym of Capensisvoluta emmanuelae (Rosso, 1985) (superseded combination)
• Athleta epigona (E. von Martens, 1904): synonym of Afriathleta epigona (E. von Martens, 1904) (unaccepted > superseded combination)
• Athleta gandilloti Chavan, 1941 †: synonym of Athleta gandilloti (Chavan, 1941) † (superseded combination)
• Athleta gilchristi (G. B. Sowerby III, 1902): synonym of Nataliavoluta gilchristi (G. B. Sowerby III, 1902) (superseded combination)
• Athleta glabrata (Kilburn, 1971): synonym of Nataliavoluta glabrata (Kilburn, 1971) (superseded combination)
• Athleta insperata Darragh, 1979: synonym of Ternivoluta insperata (Darragh, 1979) (superseded combination)
• Athleta kilburni (Rehder, 1974): synonym of Afriathleta kilburni (Rehder, 1974) (superseded combination)
• Athleta lutosa (H. J. Koch, 1948): synonym of Capensisvoluta lutosa (H. J. Koch, 1948) (superseded combination)
• Athleta massieri Petuch & Berschauer, 2017: synonym of Capensisvoluta massieri (Petuch & Berschauer, 2017)
• Athleta mozambicana (Rehder, 1972): synonym of Nataliavoluta mozambicana (Rehder, 1972) (superseded combination)
• Athleta nana (Rehder, 1974): synonym of Nataliavoluta nana (Rehder, 1974) (superseded combination)
• Athleta nicklesi (Rosso, 1976): synonym of Capensisvoluta nicklesi (Rosso, 1976)
• Athleta rarispina (Lamarck, 1811) †: synonym of Athleta ficulina (Lamarck, 1811) †
• Athleta rosavittoriae (Rehder, 1981): synonym of Afriathleta rosavittoriae (Rehder, 1981) (superseded combination)
• Athleta semirugata (Rehder & C. S. Weaver, 1974): synonym of Afriathleta semirugata (Rehder & C. S. Weaver, 1974) (superseded combination)
• Athleta spinosa (Linnaeus, 1758) †: synonym of Athleta spinosus (Linnaeus, 1758) † (incorrect grammatical agreement of specific epithet)
• Athleta studeri (E. von Martens, 1897): synonym of Ternivoluta studeri (E. von Martens, 1897) (superseded combination)
• Athleta subcrenulifera Darragh, 1971 †: synonym of Ternivoluta subcrenulifera (Darragh, 1971) † (superseded combination)

• Marwick, J. (1926). Tertiary and recent Volutidae of New Zealand. Transactions of the New Zealand Institute. 56: 259-303.
• Merle & Pacaud & Marivaux, Volutidae (Mollusca: Gastropoda) of the Lakhra Formation (Earliest Eocene, Sindh, Pakistan): systematics, biostratigraphy and paleobiogeography; Zootaxa, 3826 (1) : 101-138, fig. 1-12. , 2014
• Lozouet P. (2019). Volutidae, Volutomitridae, Marginellidae, Cystiscidae et Cancellariidae (Mollusca, Gastropoda, Neogastropoda) de l'Oligocène supérieur (Chattien) du bassin de l'Adour (Sud-Ouest de la France). Cossmanniana. 20: 3–109
• Veldsman, S.G. & Veldsman, J.H. (V.). (2022). A taxonomic revision of the Volutidae of southern Africa, including the description of three new genera and several new species. Visaya. suppl. 17: 1-168.

• Conrad, T. A. (1853). Synopsis of the genera Cassidula, Humph., and of a proposed new genus Athleta. Proceedings of the Academy of Natural Sciences of Philadelphia. 6(11-12): 448-449
• Sacco, F. (1890). Catalogo paleontologico del bacino terziaro del Piemonte. Bollettino della Società Geologica Italiana. 8(3): 281-356; 9(2): 185-340
• Newton R.B. (1906). Note on Swainson's genus Volutilithes. Proceedings of the Malacological Society of London. 7(2): 100-104, pl. 12
• Cotton, B. C. (1949). Australian Recent and Tertairy Mollusca, Family Volutidae. Records of the South Australian Museum. 9: 181-196
• Maxwell, P.A. (2003). The volutid genera Athleta and Lyria (Mollusca: Gastropoda) in the New Zealand Cenozoic. Journal of the Royal Society of New Zealand. 33: 363-394

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